Tawny Owl

识别特征

The tawny owl is a robust bird, 37–46 cm (15–18 in) in length, with an 81–105 cm (32–41 in) wingspan. Weight can range from 385 to 800 g (0.849 to 1.764 lb). Its large rounded head lacks ear tufts, and the facial disc surrounding the dark brown eyes is usually rather plain. The nominate race has two morphs which differ in their plumage colour, one form having rufous brown upperparts and the other greyish brown, although intermediates also occur. The underparts of both morphs are whitish and streaked with brown. Feathers are moulted gradually between June and December. This species is sexually dimorphic; the female is much larger than the male, 5% longer and more than 25% heavier.<p>The tawny owl flies with long glides on rounded wings, less undulating and with fewer wingbeats than other Eurasian owls, and typically at a greater height. The flight of the tawny owl is rather heavy and slow, particularly at takeoff, though the bird can attain a top flight speed of around 50 mph. As with most owls, its flight is silent because of its feathers' soft, furry upper surfaces and a fringe on the leading edge of the outer primaries. Its size, squat shape and broad wings distinguish it from other owls found within its range; the great grey owl (Strix nebulosa), Eurasian eagle-owl (Bubo bubo) and Ural owl (Strix uralensis) are similar in shape, but much larger.<p>An owl's eyes are placed at the front of the head and have a field overlap of 50–70%, giving it better binocular vision than diurnal birds of prey (overlap 30–50%). The tawny owl's retina has about 56,000 light-sensitive rod cells per square millimetre (36 million per square inch); although earlier claims that it could see in the infrared part of the spectrum have been dismissed, it is still often said to have eyesight 10 to 100 times better than humans in low-light conditions. However, the experimental basis for this claim is probably inaccurate by at least a factor of 10. The owl's actual visual acuity is only slightly greater than that of humans, and any increased sensitivity is due to optical factors rather than to greater retinal sensitivity; both humans and owl have reached the limit of resolution for the retinas of terrestrial vertebrates.<p>Adaptations to night vision include the large size of the eye, its tubular shape, large numbers of closely packed retinal rods, and an absence of cone cells, since rod cells have superior light sensitivity. There are few coloured oil drops, which would reduce the light intensity. Unlike diurnal birds of prey, owls normally have only one fovea, and that is poorly developed except in daytime hunters such as the short-eared owl.<p>Hearing is important for a nocturnal bird of prey, and as with other owls, the tawny owl's two ear openings differ in structure and are asymmetrically placed to improve directional hearing. A passage through the skull links the eardrums, and small differences in the time of arrival of a sound at each ear enables its source to be pinpointed. The left ear opening is higher on the head than the larger right ear and tilts downward, improving sensitivity to sounds from below. Both ear openings are hidden under the facial disk feathers, which are structurally specialized to be transparent to sound, and are supported by a movable fold of skin (the pre-aural flap).<p>The internal structure of the ear, which has large numbers of auditory neurons, gives an improved ability to detect low-frequency sounds at a distance, which could include rustling made by prey moving in vegetation. The tawny owl's hearing is ten times better than a human's, and it can hunt using this sense alone in the dark of a woodland on an overcast night, but the patter of raindrops makes it difficult to detect faint sounds, and prolonged wet weather can lead to starvation if the owl cannot hunt effectively.<p>The commonly heard female contact call is a shrill, kew-wick but the male has a quavering advertising song hoo...ho, ho, hoo-hoo-hoo-hoo. William Shakespeare used this owl's song in Love's Labour's Lost (Act 5, Scene 2) as "Then nightly sings the staring owl, Tu-whit; Tu-who, a merry note, While greasy Joan doth keel the pot", but this stereotypical call is actually a duet, with the female making the kew-wick sound, and the male responding hooo. The call is easily imitated by blowing into cupped hands through slightly parted thumbs, and a study in Cambridgeshire found that this mimicry produced a response from the owl within 30 minutes in 94% of trials. A male's response to a broadcast song appears to be indicative of his health and vigour; owls with higher blood parasite loads use fewer high frequencies and a more limited range of frequencies in their responses to an apparent intruder. The vocal activity of tawny owls depends on sex, annual cycle stage and weather, with males being more vocal than females year-round, with peak vocal activity during incubation and post-breeding.<p>Although both colour morphs occur in much of the European range, brown birds predominate in the more humid climate of western Europe, with the grey morph becoming more common further east; in the northernmost regions, all the owls are a cold-grey colour. The Siberian and Scandinavian subspecies are 12% larger and 40% heavier, and have 13% longer wings than western European birds, in accordance with Bergmann's rule which predicts that northern forms will typically be bigger than their southern counterparts.<p>The plumage colour is genetically controlled, and studies in Finland and Italy indicate that grey-morph tawny owls have more reproductive success, better immune resistance, and fewer parasites than brown birds. Although this might suggest that eventually the brown morph could disappear, the owls show no colour preference when choosing a mate, so the selection pressure in favour of the grey morph is reduced. There are also environmental factors involved. The Italian study showed that brown-morph birds were found in denser woodland, and in Finland, Gloger's rule would suggest that paler birds would in any case predominate in the colder climate.

分类信息

The species was first described by Linnaeus in his Systema naturae in 1758 under its current scientific name. The binomial derives from Greek strix "owl" and Italian allocco, "tawny owl" (from Latin ulucus "screech-owl").<p>The tawny owl is a member of the wood-owl genus Strix, part of the typical owl family Strigidae, which contains all species of owl other than the barn owls. Within its genus, the tawny owl's closest relatives are Hume's owl, Strix butleri, (formerly considered to be conspecific), the Himalayan owl, Strix nivicolum, (sometimes considered conspecific), its larger northern neighbour, the Ural owl, S. uralensis, and the North American barred owl, S. varia. The Early–Middle Pleistocene Strix intermedia is sometimes considered a paleosubspecies of the tawny owl, which would make it that species' immediate ancestor.<p>The tawny owl subspecies are often poorly differentiated, and may be at a flexible stage of subspecies formation with features related to the ambient temperature, the colour tone of the local habitat, and the size of available prey. Consequently, various authors have historically described between 10 and 15 subspecies. The seven currently recognised subspecies are listed below.

学名

Strix aluco

命名信息

Linnaeus, 1758

目

Strigiformes

科

Strigidae

属

Strix

分布与栖息地

The tawny owl is non-migratory and has a distribution stretching discontinuously across temperate Europe from Great Britain and the Iberian Peninsula eastwards to western Siberia. It is absent from Ireland - probably because of competition from the long-eared owl (Asio otus) - and only a rare vagrant to the Balearic and Canary Islands. In the Himalayas and East Asia it is replaced by the Himalayan owl (Strix nivicolum) and in northwest Africa it is replaced by the closely related Maghreb owl (Strix mauritanica).<p>This species is found in deciduous and mixed forests, and sometimes mature conifer plantations, preferring locations with access to water. Cemeteries, gardens and parks have allowed it to spread into urban areas, including central London. Although tawny owls occur in urban environments, especially those with natural forests and wooded habitat patches, they are less likely to occur at sites with high noise levels at night. The tawny owl is mainly a lowland bird in the colder parts of its range, but breeds to 550 metres (1,800 ft) in Scotland, 1,600 m (5,200 ft) in the Alps, 2,350 m (7,710 ft) in Turkey, and up to 2,800 m (9,200 ft) in Myanmar.<p>The tawny owl has a geographical range of at least 10 million km2 (3.8 million mi2) and a large population including an estimated 970,000–2,000,000 individuals in Europe alone. Population trends have not been quantified, but there is evidence of an overall increase. This owl is not believed to meet the IUCN Red List criterion of declining more than 30% in ten years or three generations and is therefore evaluated as being of least concern. In the UK it is on the RSPB Amber List of Concern. This species has expanded its range in Belgium, the Netherlands, Norway and Ukraine, and populations are stable or increasing in most European countries. Declines have occurred in Finland, Estonia, Italy and Albania. Tawny owls are listed in Appendix II of the Convention on International Trade in Endangered Species (CITES) meaning international trade (including in parts and derivatives) is regulated.

迁徙

Resident species

行为

Migration pattern: Resident species.

保护状况

Population trend: stable.

种群趋势

stable