Râle des genêts

Identification

The corn crake is a medium-sized rail, 27–30 cm (11–12 in) long with a wingspan of 42–53 cm (17–21 in). Males weigh 165 g (5.8 oz) on average and females 145 g (5.1 oz). The adult male has the crown of its head and all of its upperparts brown-black in colour, streaked with buff or grey. The wing coverts are a distinctive chestnut colour with some white bars. The face, neck and breast are blue-grey, apart from a pale brown streak from the base of the bill to behind the eye, the belly is white, and the flanks, and undertail are barred with chestnut and white. The strong bill is flesh-coloured, the iris is pale brown, and the legs and feet are pale grey. Compared to the male, the female has warmer-toned upperparts and a narrower duller eye streak. Outside the breeding season, the upperparts of both sexes become darker and the underparts less grey. The juvenile is like the adult in appearance, but has a yellow tone to its upperparts, and the grey of the underparts is replaced with buff-brown. The chicks have black down, as with all rails. While there are no subspecies, all populations show great individual variation in colouring, and the birds gradually become paler and greyer towards the east of the range. Adults undergo a complete moult after breeding, which is normally finished by late August or early September, before migration to south eastern Africa. There is a pre-breeding partial moult prior to the return from Africa, mainly involving the plumage of the head, body and tail. Young birds have a head and body moult about five weeks after hatching.<p>The corn crake is sympatric with the African crake on the wintering grounds, but can be distinguished by its larger size, paler upperparts, tawny upperwing and different underparts pattern. In flight, it has longer, less rounded wings, and shallower wingbeats than its African relative, and shows a white leading edge to the inner wing. In both the breeding and wintering ranges it is unlikely to be confused with any other rails, since sympatric species are smaller, with white markings on the upperparts, different underparts patterns and shorter bills. A flying corn crake can resemble a gamebird, but its chestnut wing pattern and dangling legs are diagnostic.<p>On the breeding grounds, the male corn crake's advertising call is a loud, repetitive, grating krek krek normally delivered from a low perch with the bird's head and neck almost vertical and its bill wide open. The call can be heard from 1.5 km (0.93 mi) away, and serves to establish the breeding territory, attract females, and challenge intruding males. Slight differences in vocalisations mean that individual males can be distinguished by their calls. Early in the season, the call is given almost continuously at night, and often during the day, too. It may be repeated more than 20,000 times a night, with a peak between midnight and 3 am. The call has evolved to make a singing male's location clear, as this species hides in vegetation. The frequency of calling reduces after a few weeks but may intensify again near the end of the laying period before falling away towards the end of the breeding season. To attract males, mechanical imitations of their call can be produced by rubbing two pieces of wood or ribs, one of them with notches, or by flicking a credit card against a comb or zip-fastener. The male also has a growling call, given with the bill shut and used during aggressive interactions.<p>The female corn crake may give a call that is similar to that of the male; it also has a distinctive barking sound, similar in rhythm to the main call but without the grating quality. The female also has a high-pitched cheep call, and a oo-oo-oo sound to call the chick. The chicks make a quiet peeick-peeick contact call, and a chirp used to beg for food. Because of the difficulty in seeing this species, it is usually censused by counting males calling between 11 pm and 3 am; the birds do not move much at night, whereas they may wander up to 600 m (660 yd) during the day, which could lead to double-counting if monitored then. Identifying individual males suggests that just counting calling birds underestimates the true count by nearly 30%, and the discrepancy is likely to be greater, since only 80% of males may call at all on a given night. The corn crake is silent in Africa.

Taxonomie

The rails are a bird family comprising nearly 150 species. Although the origins of the group are lost in antiquity, the largest number of species and least specialised forms are found in the Old World, suggesting this family originated there. The taxonomy of the small crakes is complicated, but the closest relative of the corn crake is the African crake, which has been given its own genus, Crecopsis.<p>Corn crakes were first described by Carl Linnaeus in his 1758 10th edition of Systema Naturae as Rallus crex, but was subsequently moved to the genus Crex, created by German naturalist and ornithologist Johann Matthäus Bechstein in 1803, and named Crex pratensis. The earlier use of crex gives it priority over Bechstein's specific name pratensis, and leads to the current name of Crex crex. The binomial name, Crex crex, from the ancient Greek "κρεξ", is onomatopoeic, referring to the crake's repetitive grating call. The common name was formerly spelt as a single word, "corncrake", but the official version is now "corn crake". The English names refer to the species' habit of nesting in dry hay or cereal fields, rather than marshes used by most members of this family.

Nom scientifique

Crex crex

Autorité

(Linnaeus, 1758)

Ordre

Gruiformes

Famille

Rallidés

Genre

Crex

Répartition et habitat

The corn crake breeds from Ireland east through Europe to central Siberia. Although it has vanished from much of its historic range, this bird was once found in suitable habitats in Eurasia everywhere between latitudes 41°N and 62°N. There is also a sizable population in western China, but this species nests only rarely in northern Spain and in Turkey. Old claims of breeding in South Africa are incorrect, and result from misidentification of eggs in a museum collection which are actually those of the African rail.<p>The corn crake winters mainly in Africa, from the Democratic Republic of the Congo and central Tanzania south to eastern South Africa. North of this area, it is mainly seen on migration, but occasionally winters in North Africa and to the west and north of its core area in southeast Africa. Most of the South African population of about 2,000 birds occurs in KwaZulu-Natal and the former Transvaal Province, and numbers elsewhere in Africa are uncertain. There are several nineteenth-century records, when populations were much higher than now, of birds being seen in western Europe, mainly Britain and Ireland, between December and February.<p>This crake migrates to Africa along two main routes: a western route through Morocco and Algeria, and a more important flyway through Egypt. On passage, it has been recorded in most countries between its breeding and wintering ranges, including much of West Africa. Birds from Coll following the western route paused in West Africa on their way further south, and again on the return flight, when they also rested in Spain or North Africa. Eastern migrants have been recorded in those parts of southern Asia that lie between the east of the breeding range and Africa. Further afield, the corn crake has been recorded as a vagrant to Sri Lanka, Vietnam and Australia, New Zealand, the Seychelles, Bermuda, Canada, the US, Greenland, Iceland, the Faroes, the Azores, Madeira, and the Canary Islands.<p>The corn crake is mainly a lowland species, but breeds up to 1,400 m (4,600 ft) altitude in the Alps, 2,700 m (8,900 ft) in China and 3,000 m (9,800 ft) in Russia. When breeding in Eurasia, the corn crake's habitats would originally have included river meadows with tall grass and meadow plants including sedges and irises. It is now mainly found in cool moist grassland used for the production of hay, particularly moist traditional farmland with limited cutting or fertiliser use. It also utilises other treeless grasslands in mountains or taiga, on coasts, or where created by fire. Moister areas like wetland edges may be used, but very wet habitats are avoided, as are open areas and those with vegetation more than 50 cm (20 in) tall, or too dense to walk through. The odd bush or hedge may be used as a calling post. Grassland which is not mown or grazed becomes too matted to be suitable for nesting, but locally-grown crops such as cereals, peas, rape, clover or potatoes may be used. After breeding, adults move to taller vegetation such as common reed, iris, or nettles to moult, returning to the hay and silage meadows for the second brood. In China, flax is also used for nest sites. Although males often sing in intensively managed grass or cereal crops, successful breeding is uncommon, and nests in the field margins or nearby fallow ground are more likely to succeed.<p>When wintering in Africa, the corn crake occupies dry grassland and savanna habitats, occurring in vegetation 30–200 cm (0.98–6.56 ft) tall, including seasonally burnt areas and occasionally sedges or reed beds. It is also found on fallow and abandoned fields, uncut grass on airfields, and the edges of crops. It occurs at up to at least 1,750 m (5,740 ft) altitude in South Africa. Each bird stays within a fairly small area. Although it sometimes occurs with the African crake, that species normally prefers moister and shorter grassland habitats than does the corn crake. On migration, the corn crake may also occur in wheatfields and around golf courses.

Aire de reproduction

PAL : Europe to c Siberia and nw China

Aire hors reproduction

s AF

Migration

Complete migrants

Comportement

The corn crake is a difficult bird to see in its breeding sites, usually being hidden by vegetation, but will sometimes emerge into the open. Occasionally, individuals may become very trusting; for five consecutive summers, an individual crake on the Scottish island of Tiree entered a kitchen to feed on scraps, and, in 1999, a wintering Barra bird would come for poultry feed once the chickens had finished. In Africa, it is more secretive than the African crake, and, unlike its relative, it is rarely seen in the open, although it occasionally feeds on tracks or road sides. The corn crake is most active early and late in the day, after heavy rain and during light rain. Its typical flight is weak and fluttering, although less so than that of the African crake. For longer flights, such as migration, it has a steadier, stronger action with legs drawn up. It walks with a high-stepping action, and can run swiftly through grass with its body held horizontal and laterally flattened. It will swim if essential. When flushed by a dog, it will fly less than 50 m (160 ft), frequently landing behind a bush or thicket, and then crouch on landing. If disturbed in the open, this crake will often run in a crouch for a short distance, with its neck stretched forward, then stand upright to watch the intruder. When captured it may feign death, recovering at once if it sees a way out.<p>The corn crake is solitary on the wintering grounds, where each bird occupies 4.2–4.9 ha (10–12 acres) at one time, although the total area used may be double that, since an individual may move locally due to flooding, plant growth, or grass cutting. Flocks of up to 40 birds may form on migration, sometimes associating with common quails. Migration takes place at night, and flocks resting during the day may aggregate to hundreds of birds at favoured sites. The ability to migrate is innate, not learned from adults. Chicks raised from birds kept in captivity for ten generations were able to migrate to Africa and return with similar success to wild-bred young.<p>Until 1995, it was assumed that the corn crake is monogamous, but it transpires that a male may have a shifting home range, and mate with two or more females, moving on when laying is almost complete. The male's territory can vary from 3 to 51 ha (7.4 to 126.0 acres), but averages 15.7 ha (39 acres). The female has a much smaller range, averaging only 5.5 ha (14 acres). A male will challenge an intruder by calling with his wings drooped and his head pointing forward. Usually the stranger moves off; if it stays, the two birds square up with heads and necks raised and the wings touching the ground. They then run around giving the growling call and lunging at each other. A real fight may ensue, with the birds leaping at each other and pecking, and sometimes kicking. Females play no part in defending the territory.<p>The female may be offered food by the male during courtship. He has a brief courtship display in which the neck is extended and the head held down, the tail is fanned, and the wings are spread with the tips touching the ground. He will then attempt to approach the female from behind, and then leap on her back to copulate. The nest is typically in grassland, sometimes in safer sites along a hedge, or near an isolated tree or bush, or in overgrown vegetation. Where grass is not tall enough at the start of the season, the first nest may be constructed in herby or marsh vegetation, with the second brood in hay. The second nest may also be at a higher altitude that the first, to take advantage of the later-developing grasses further up a hill. The nest, well hidden in the grass, is built in a scrape or hollow in the ground. It is made of woven coarse dry grass and other plants, and lined with finer grasses. Although nest construction is usually described as undertaken by the female, a recent aviary study found that in the captive population the male always built the nest.<p>The nest is 12–15 cm (4.7–5.9 in) in diameter and 3–4 cm (1.2–1.6 in) deep. The clutch is 6–14, usually 8–12 eggs; these are oval, slightly glossy, creamy or tinted with green, blue or grey, and blotched red-brown. They average 37 mm × 26 mm (1.5 in × 1.0 in) and weigh about 13–16 g (0.46–0.56 oz), of which 7% is shell. The eggs are laid at daily intervals, but second clutches may sometimes have two eggs added per day. Incubation is by the female only; her tendency to sit tight when disturbed, or wait until the last moment to flee, leads to many deaths during hay-cutting and harvesting. The eggs hatch together after 19–20 days, and the precocial chicks leave the nest within a day or two. They are fed by the female for three or four days, but can find their own food thereafter. The juveniles fledge after 34–38 days. The second brood is started about 42 days after the first, and the incubation period is slightly shorter at 16–18 days. The grown young may stay with the female until departure for Africa.<p>Nest success in undisturbed sites is high, at 80–90%, but much lower in fertilised meadows and on arable land. The method and timing of mowing is crucial; mechanized mowing can kill 38–95% of chicks in a given site, and losses average 50% of first brood chicks and somewhat less than 40% of second brood chicks. The influence of weather on chick survival is limited; although chick growth is faster in dry or warm weather, the effects are relatively small. Unlike many precocial species, chicks are fed by their mother to a greater or lesser extent until they become independent, and this may cushion them from adverse conditions. The number of live chicks hatched is more important than the weather, with lower survival in large broods. The annual adult survival rate is under 30%, although some individuals may live for 5–7 years.<p>The corn crake is omnivorous, but mainly feeds on invertebrates, including earthworms, slugs and snails, spiders, beetles, dragonflies, grasshoppers and other insects. In the breeding areas, it is a predator of Sitona weevils, which infest legume crops. and in the past consumed large amounts of the former grassland pests, leatherjackets and wireworms. This crake will also eat small frogs and mammals, and plant material including grass seed and cereal grain. Its diet on the wintering grounds is generally similar, but includes locally available items such as termites, cockroaches and dung beetles. Food is taken from the ground, low-growing plants and from inside grass tussocks; the crake may search leaf litter with its bill, and run in pursuit of active prey. Hunting is normally in cover, but, particularly in the wintering areas, it will occasionally feed on grassy tracks or dirt roads. Indigestible material is regurgitated as 1 cm (0.39 in) pellets. Chicks are fed mainly on animal food, and when fully grown they may fly with the parents up to 6.4 km (4.0 mi) to visit supplementary feeding areas. As with other rails, grit is swallowed to help break up food in the stomach.

Conservation

Tendance de la population

stable