Arctic Loon

Identification

The adult black-throated loon is 58 to 73 cm (23 to 29 in) in length with a 100 to 130 cm (39 to 51 in) wingspan and a weight of 1.3 to 3.4 kilograms (2.9 to 7.5 lb). The nominate subspecies in its breeding plumage has a grey head and hindneck, with a black throat and a large black patch on the foreneck, both of which have a soft purple gloss. The lower throat has a necklace-shaped patch of short parallel white lines. The sides of the throat have about five long parallel white lines that start at the side of the patch on the lower throat and run down to the chest, which also has a pattern of parallel white and black lines. The rest of the underparts, including the centre of the chest, are pure white. The upperparts are blackish down to the base of the wing, where there are a few rows of high contrast white squares that cover the mantle and scapulars. There are small white spots on both the lesser and median coverts. The rest of the upperwing is a blackish colour. The underwing is paler than the upperwing, and the underwing coverts are white. The tail is blackish. The bill and legs are black, with a pale grey colour on the inner half of the legs. The toes and the webs are grey, the latter also being flesh coloured. The irides are a deep brown-red. The sexes are alike, and the subspecies viridigularis is very similar to the nominate except that the former has a green throat patch, instead of black. The subspecies viridigularis does still retain a purplish gloss, although it is less than the nominate.<p>The non-breeding adult differs from the breeding adult in that the cap and the back of the neck are more brownish. The non-breeding adult also lacks the patterned upperparts of the breeding adult, although some of the upperwing coverts do not lose their white spots. This results in the upperparts being an almost unpatterned black from above. The sides of the throat are usually darker at the white border separating the sides of the throat and the front of the throat; most of the time a thin dark necklace between these two areas can be seen. There is white on the sides of the head that are below the eye. The bill is a steel-grey with, similar to the breeding adult, a blackish tip.<p>The juvenile is similar to the non-breeding adult, but has a browner appearance. It has a buffy scaling on the upperparts that is especially pronounced on the scapulars. The lower face and front of the neck has a diffused brownish tinge. The juvenile does not have the white spots on the wing coverts, and its irides are darker and more dull in colour. The chick hatches with down feathers that range in colour from sooty-brown to brownish-grey, usually with a slightly paler head. The abdomen is pale.<p>The black-throated loon can be distinguished from the Pacific loon by the white on the flanks of the former.<p>The male, when breeding, vocalizes a loud and rhythmic "oooéé-cu-cloooéé-cu-cloooéé-cu-cluuéé" whistling song. A "áááh-oo" wail can also be heard, and a growling or croaking "knarr-knor", a sound given especially at night. The alarm call at the nest is a rising "uweek".

Taxonomy

The black-throated loon, Gavia arctica, was originally described by Carl Linnaeus as Colymbus arcticus in his 18th-century work, Systema Naturae. It was moved to the genus Gavia by the American Ornithologists' Union (AOU) in 1897. The genus name Gavia comes from the Latin for "sea mew", as used by the ancient Roman naturalist Pliny the Elder. The specific arctica is Latin for "northern" or "Arctic". The name of the subspecies viridigularis stems from the Latin "viridis", meaning "green", and the Latin "gularis", meaning "throated", in reference to the green throat of this subspecies. The common name, black-throated loon, stems from its black throat patch. This loon is also called the Arctic loon and the black-throated diver.<p>There are two subspecies:<p>G. a. viridigularis was considered to be a separate species when described by Jonathan Dwight in 1918, but a year after, in 1919, Arthur Cleveland Bent suggested that the former species be moved to its current placement as a subspecies. The black-throated loon was previously considered conspecific with the North American Pacific loon, which was its subspecies, but they have now been split into two species; there was no evidence of the two interbreeding in areas where they occurred together. Furthermore, the architecture of the air sacs in the lungs of the two species are significantly different. This split was done by the AOU in 1985. The phylogeny of this species is debated, the black-throated loon and the Pacific loon traditionally being considered sister species, whereas a study using mitochondrial and nuclear intron DNA supported placing the black-throated loon sister to a clade consisting of the Pacific loon and the two sister species that are the common loon and the yellow-billed loon. This latter study is criticized on the basis that it may form a phylogeny on incomplete lineage sorting. In the former phylogeny, the split between the Pacific loon and the black-throated loon is proposed to have happened about 6.5 million years ago.

Scientific name

Gavia arctica

Authority

(Linnaeus, 1758)

Order

Gaviiformes

Family

Gaviidae

Genus

Gavia

Range and habitat

The black-throated loon has a large range, breeding taking place across northern Europe, Asia, and the Seward Peninsula in Alaska. When breeding, it is found in the area around isolated, deep freshwater lakes larger than 0.1 square kilometres (0.039 sq mi), especially those with inlets, as it prefers to face only small stretches of open water. When it is not breeding, this loon moves in a general southward direction and towards ice-free sea, usually wintering in coasts on north-east Atlantic Ocean and those on the eastern and western Pacific Ocean, such as the coasts of Japan. During this time, its habitat is usually inshore waters along sheltered coasts, although it will sometimes be found inland, in places such as the Mediterranean and Black seas.

Breeding range

PAL, NA : Arctic to temperate n, c Eurasia and far w Alaska

Non-breeding range

to coastal s European and w Asia seas; to Japan and Korean Pen.

Migration

Complete migrants

Behavior

Migration pattern: Complete migrants.

Conservation

Population trend: declining.

Population trend

declining