Saxaul Sparrow

Identification

The saxaul sparrow is one of the larger sparrows at 14–16 centimetres (5.5–6.3 in) and 25–32 grams (0.88–1.13 oz). Wing length varies from 7.1 to 8.1 centimetres (2.8 to 3.2 in), with males generally being larger. The tail is short at 6.3–6.95 centimetres (2.48–2.74 in). The saxaul sparrow's legs are pale or pinkish brown, with a tarsus length of 1.95 inches (50 mm). Its bill is 1.0–1.3 centimetres (0.39–0.51 in) long, pale grey on the juvenile, pale yellowish with a black tip on the breeding female, and black on the breeding male. Like all other sparrows, it flies swiftly and often at height.<p>Distinctive markings, especially on its head, make the saxaul sparrow unlikely to be confused with any other bird. It is dull-coloured, with plumage ranging between dull grey and warm sandy brown, varying between and within subspecies. Birds of the subspecies ammodendri are a sandy grey, while nigricans birds are similar but darker, and stoliczkae birds are warm brown or russet. Birds of the subspecies stoliczkae and those from the southwest of the range of ammodendri also differ from usual ammodendri birds in their lack of streaking on the rump and upper tail coverts. Birds in Mongolia have a larger and deeper bill and broad bluish streaks on their chest.<p>The male saxaul sparrow has bold markings, with a black stripe along the top of its head and another through its eye. It has black feathering, or a "bib", on its throat and upper belly. By comparison to other sparrows this is thin on the throat, but wide on the breast. The male has a bright russet patch on the sides of its crown and nape. Its cheeks are pale grey or buff, and its underparts are whitish, tinged buff or grey on its sides. Its back is grey or warm brown, streaked variably with black. Its shoulders are more lightly streaked with black bars. The male's thin tail is brown, with the edges and tips of feathers paler. Its median coverts are black with a white tip, while its other wing feathers are variably dark brown, cinnamon, or black, tipped buff or whitish and edged grey. The non-breeding male differs in having slightly paler plumage.<p>The female is similar in some ways to the male, but paler and duller. It is sandy grey or brown, with a back patterned like that of the male, and white or whitish underparts. The head of the females of the subspecies ammodendri and nigricans is dingy grey with darker smudges on the forehead, behind its eyes, and on its throat. The female of the subspecies stoliczkae is buff-brown with a white throat, a conspicuous pale supercilium, darker forehead, and lighter cheeks. The juvenile is similar to the female, differing in its lack of dark tinges on its throat and crown. In adults, moulting begins in July and ends in late August or early September. The post-juvenile moult is complete, and occurs variously from June to August.<p>The saxaul sparrow's vocalisations are little reported. Its common call is a chirp, transcribed as cheerp cheerp, softer and more melodious than that of the house sparrow. It gives a flight call transcribed as twerp, and a song described by Russian naturalist V. N. Shnitnikov as "not loud, but pleasantly melodious with fairly diversified intonations".

Taxonomy

The saxaul sparrow was first described by English zoologist John Gould in a March 1872 instalment of The Birds of Asia, from a specimen collected near Kyzylorda, now in southern Kazakhstan, by Russian naturalist Nikolai Severtzov. Severtzov had been planning to describe the species as Passer ammodendri for several years and had been distributing specimens among other naturalists. When natural history dealer Charles Dode escaped from the Paris Commune in 1871 with some of his collection, Gould obtained specimens from a set of rare birds Dode exhibited to the Zoological Society of London. Severtzov did not describe the species until 1873, and some later writers preferred to attribute him, but Gould's description takes priority over Severtzov's. The saxaul sparrow's species name refers to its desert habitat, coming from the name of the Ammodendron or sand acacia tree, which is in turn derived from the Ancient Greek άμμος (ammos, "sand") and δένδρον (dendron, "tree"). The English name saxaul sparrow refers to the saxaul plant, with which it is closely associated. The saxaul sparrow usually is classified in the genus Passer with the house sparrow and around twenty other species, although a genus Ammopasser was created for the saxaul sparrow by Nikolai Zarudny in 1890.<p>The saxaul sparrow's relations within the genus Passer are unclear, although with its black throat feathering it has usually been considered part of the "Palaearctic black-bibbed sparrow" group related to the house sparrow. J. Denis Summers-Smith considered that the Palaearctic Passer sparrows evolved about 25,000 to 15,000 years ago, during the last glacial period. During this time, sparrows would have been isolated in ice-free refugia, such as a certain steppe region in Central Asia, where Summers-Smith suggested the saxaul sparrow evolved. Genetic and fossil evidence suggest a much earlier origin for the Passer species, perhaps in the Miocene and Pliocene, as suggested by Luis Allende and colleagues in their 2001 phylogenetic analysis of mitochondrial DNA. This analysis also suggested that the saxaul sparrow may be an early offshoot or basal species in its genus, a relative of certain African sparrows such as the northern grey-headed sparrow. If the saxaul sparrow is related to these species, either the saxaul sparrow formerly occurred in the deserts of Africa and Arabia, or each of the groups of Passer sparrows are of African origin.<p>Across its Central Asian distribution, the saxaul sparrow occurs in six probably disjunct areas, and is divided into at least three subspecies. The nominate subspecies Passer ammodendri ammodendri inhabits three of these areas, one in the Syr Darya basin of Kazakhstan and Uzbekistan, and another to the south of Lake Balkhash and the north of Almaty, where it is only common in the valley of the Ili River. In a third area, sometimes recognised as a subspecies korejewi, ammodendri birds breed sporadically in parts of central Turkmenistan, Iran, and possibly Afghanistan, migrating to the south during the winter. The subspecies stoliczkae was named after Ferdinand Stoliczka in 1874 by Allan Octavian Hume, from specimens Stoliczka collected in Yarkand. This subspecies is separated from the other two subspecies by the Tian Shan mountains. It is found across a broad swath of China from Kashgar east to the far west of Inner Mongolia, through the areas around the Taklamakan Desert (but probably not in the inhospitable desert itself), and through the east of Xinjiang, northern Gansu, and the fringes of southern Mongolia. In the extreme west of the Gobi Desert a disjunct population separated from the other stoliczkae birds by the Gurvan Saikhan Uul mountains occurs, which is sometimes separated as a subspecies timidus. The subspecies nigricans, described by ornithologist L. S. Stepanyan in 1961, is found in northern Xinjiang's Manasi River valley.

Scientific name

Passer ammodendri

Authority

Gould, 1872

Order

Passeriformes

Family

Passeridae

Genus

Passer

Range and habitat

The saxaul sparrow is found in remote parts of Central Asia, where its distribution is believed to fall into six disjunct areas, although this is uncertain due to the scarcity of records. It is found in deserts, especially around rivers and oases. It is usually found around shrubs such as saxaul (Haloxylon), poplar (Populus), or tamarisk (Tamarix). Sometimes it occurs around settlements and grain fields, especially during the winter. It is not believed to be threatened, since it is reported as locally common across a wide range, and hence it is assessed as Least Concern on the IUCN Red List. However, it seems to have lost large parts of its range to the intensification of agriculture and desertification caused by overgrazing.

Migration

Resident species

Behavior

Migration pattern: Resident species.

Conservation

Population trend: stable.

Population trend

stable