Yellowhammer

Identification

The yellowhammer is a large bunting, 16–16.5 cm (6.3–6.5 in) long, with a 23–29.5 cm (9.1–11.6 in) wingspan; it weighs 20–36.5 g (0.71–1.29 oz). The male of the nominate subspecies E. c. citrinella has a bright yellow head, heavily streaked brown back, rufous rump, yellow under parts, and white outer tail feathers. The female is less brightly coloured, and more streaked on the crown, breast, and flanks. Both sexes are less strongly marked outside the breeding season, when the dark fringes on new feathers obscure the yellow plumage. The juvenile is much duller and less yellow than the adults, and often has a paler rump.<p>After breeding, adults have a complete moult, which takes at least eight weeks; males acquire more yellow in the plumage each time they moult. Juveniles have a partial moult not long after fledging, replacing the head, body, and some covert feathers.<p>Differences between the subspecies are small and geographically gradual. On average, the male of E. c. caliginosa is slightly smaller and darker than the same sex of the nominate subspecies, and also has more streaking on its back, a greenish tint to the yellow of the head and more chestnut on the flanks. The male of the eastern form, E. c. erythrogenys, is paler and less streaked than E. c. citrinella. Its flanks, undertail and wing bars are usually whiter, and its crown and throat are brighter yellow. Distinguishing females of the three subspecies using plumage features is not usually possible.<p>Females and juveniles, especially of the pale eastern subspecies, E. c. erythrogenys, may be confused with pine buntings, but they always have a yellow tint to their plumage, a paler rufous rump, and more uniform upperparts than that species. Young and female yellowhammers can be distinguished from cirl buntings by the grey-brown rump of the latter species. Male hybrids with pine buntings are typically white-faced and have some yellow on the head, under parts or flight feathers, but females are usually indistinguishable from yellowhammers.<p>The song of the cock yellowhammer is a series of short notes, gradually increasing in volume and followed by one or two more protracted notes. It is often represented as "A little bit of bread and no cheese", and the full version can be confused with the almost identical song of the pine bunting. If the final notes are omitted, confusion with the cirl bunting is possible. Other vocalisations include a zit contact call, a see alarm, and a trilled tirrr given in flight.<p>Yellowhammer males learn their songs from their fathers, and over time, regional dialects have developed, with minor differences to the conclusion of the basic song; all are mutually recognised by birds from different areas. Each male has an individual repertoire of song variants within its regional dialect; females tend to mate with males that share their dialect, and prefer those with the largest repertoires.<p>The pine bunting and yellowhammer are so closely related that each responds to the other's song. The male yellowhammer's song is more attractive to females, and is one reason for the dominance of that species where the ranges overlap.

Taxonomy

The yellowhammer was described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae under its current scientific name. Emberiza is derived from the Old German Embritz, a bunting, and citrinella is the Italian for a small yellow bird. The English name is thought to have come from Ammer, another German word for a bunting, and was first recorded in 1553 as yelambre.<p>The bird family Emberizidae contains a single genus Emberiza, with around 40 members, that are confined to the Old World. Within its genus, the yellowhammer is most closely related to the pine bunting, with which it forms a superspecies; they have at times been considered as one species. The white-capped and cirl buntings are also near relatives of the species pair. Where their ranges meet, the yellowhammer and pine bunting interbreed; the yellowhammer is dominant, and the hybrid zone is moving further east.<p>There are currently 3 recognised subspecies of yellowhammer:

Scientific name

Emberiza citrinella

Authority

Linnaeus, 1758

Order

Passeriformes

Family

Emberizidae

Genus

Emberiza

Range and habitat

The yellowhammer breeds across the Palearctic between the 16–20 °C (61–68 °F) July isotherms. It is the commonest and most widespread European bunting, although it is absent from high mountains, Arctic regions, the western Netherlands, most of Iberia and Greece, and low-lying regions of other countries adjoining the Mediterranean Sea. It breeds in Russia east to Irkutsk, and in most of Ukraine. The Asian range extends into northwest Turkey, the Caucasus, and northern Kazakhstan.<p>Most European yellowhammers winter within their breeding range, only the far north being vacated, although some birds move south of their breeding range in Spain, Italy, and other Mediterranean countries. Distances travelled can be up to 500 km (310 mi) for northern birds. Asian birds are more strongly migratory, deserting much of the north to winter in Iraq, Iran, and southern Central Asia. The yellowhammer has occurred as a vagrant in the United Arab Emirates, Egypt, Kuwait, Morocco, Malta, the Himalayas (winter vagrant from northern Afghanistan to central Nepal), the Balearic Islands, Iceland, and the Faroes.<p>Yellowhammers of the British and Irish race, E. c. caliginosa, were introduced to New Zealand by local acclimatisation societies in 1862, and soon spread over the main islands. They sometimes visit New Zealand's subantarctic islands, although rarely staying to breed, and have reached Australia's Lord Howe Island on a number of occasions. At the beginning of the 20th century, this bunting was seen as a serious agricultural pest in its adopted country.<p>Populations of yellowhammer have also been introduced to the Falkland Islands and South Africa.<p>The yellowhammer is a bird of dry, open country, preferably with a range of vegetation types and some trees from which to sing. It is absent from urban areas, forests, and wetlands. Probably originally found at forest edges and large clearing, it has benefited from traditional agriculture, which created extensive open areas with hedges and clumps of trees.

Breeding range

PAL : subarctic to s temperate montane w, c Palearctic edge and openings: British Is., n Scandinavia to n Iberian Pen., e through subarctic Russia to c Siberia, s through montane n Mediterranean region, Balkans and Caucasus, e through n Kazakhstan and far nc Mongolia

Non-breeding range

short-range to moderate southward movement to fields, orchards: to s Iberian Pen., Turkey, Israel, w Iran, s Kazakhstan to n Afghanistan

Migration

Complete migrants

Behavior

Breeding normally starts in early May, but often in April in the south of the range. Yellowhammers are monogamous and breed when aged one year. The males establish territories along hedges or woodland fringes and sing from a tree or bush, often continuing well into July or August. The male displays to the female by raising his wings and running towards her. The nest is built by the female on or near the ground, and is typically well hidden in tussocks, against a bank or low in a bush. It is constructed from nearby plant material, such as leaves, dry grass, and stalks, and is lined with fine grasses and sometimes animal hair. It is 11.5–13 cm (4.5–5.1 in) across with a cup 4–4.5 cm (1.6–1.8 in) deep.<p>The clutch is usually three to five whitish eggs, typically patterned with a network of fine, dark lines. The eggs average 21 mm × 16 mm (0.83 in × 0.63 in) in size and weigh 2.9 g (0.10 oz), of which 6% is shell. The female incubates the eggs for 12–14 days to hatching, and broods the altricial, downy chicks until they fledge 11–13 days later. Both adults feed the chick in the nest and two or three broods are raised each year.<p>The adult annual survival rate in the UK is around 54%, and that for juveniles in their first year is 53%. The typical lifespan is three years, although records from Great Britain and Germany indicate birds surviving more than 13 years.<p>Foraging is mainly on the ground, and the bird's diet consists mainly of seeds. Oily seeds, such as those of brassicas, are ignored in favour of more starchy items. Typical food plants include common nettle, docks, common knotgrass, fat hen, common chickweed, and yarrow. Grasses are also important, particularly cereals, and grain makes up a significant part of the food consumed in autumn and winter, wheat and oats being preferred to barley. When not breeding, yellowhammers forage in flocks that can occasionally number hundreds of birds, and often contain other buntings and finches.<p>The yellowhammer adds invertebrates to its diet in the breeding season, particularly as food for its growing chicks. A wide range of species is taken, including springtails, grasshoppers, flies, beetles, caterpillars, earthworms, spiders, and snails. During the first few days, chicks are exclusively fed invertebrate prey, but from day three they are also fed cereal grains, which the chicks can digest efficiently. This is thought to be intentional by the parents to allow the nestlings to adjust their physiology to eating seed.

Conservation

The International Union for Conservation of Nature estimates the European population of the yellowhammer to be from 54–93 million individuals, suggesting a Eurasian total of 73–186 million birds. Although the population appears to be in a decline, the decrease is not rapid enough to trigger their vulnerability criteria. The large numbers and huge breeding range of about 12.9 million km2 (5 million sq mi), mean that this bunting is classified by the IUCN as being of least concern.<p>Populations have declined in recent decades in western Europe, including the British Isles, Belgium, the Netherlands, Austria, and Italy. The yellowhammer is a red-list (severely declining) species in Ireland and the UK. In 2016 the species went extinct on the Isle of Man. In eastern Europe, numbers appear to be stable, although the trend in Russia is unknown. Changes to agricultural practices are thought to be responsible for reduced breeding densities. The introduced population in New Zealand has been very successful, with breeding densities much higher than in the UK.

Population trend

declining